Hibiki Tachibana, Tsubasa Kazanari, and Chris Yukine work under Genjuurou Kazanari's command following the events of Symphogear G. Meanwhile, Maria Cadenzavna Eve, Kirika Akatsuki, and Shirabe Tsukuyomi are put under protective custody. The return to peace is short-lived, as Carol Malus Dienheim, an alchemist, and her group, the Autoscorers, appear, threatening to disrupt the status quo.